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Cascabel vs Ancho

The short answer

Both are dried Mexican chillies with low heat used for body rather than fire, and they are different peppers with different jobs. Cascabel is a small round chilli — the name means rattle, from the loose seeds — and is nutty and slightly smoky with a woody edge. Ancho is a dried poblano, wide and flat, and is sweet and raisin-like. Ancho supplies sweetness and cascabel supplies a savoury depth.

Side by side

Cascabel and Ancho compared side by side
 CascabelAncho
Botanical identityCapsicum annuum L.Capsicum annuum L.
Part usedfruitfruit
OriginMexico, with production concentrated in the central and northern states.Mexico, with the poblano cultivar associated with Puebla — which is where the name comes from.
Flavour notesnutty, woody, smoky, faintly tannicraisin, prune, sweet, cocoa, tobacco
Heat1500–4000 SHU (typically 2500 SHU)1000–2000 SHU (typically 1500 SHU)
FlavourNutty, woody and faintly tannic, with a clean moderate heat and very little sweetness. It is the least fruity of the standard Mexican dried chillies, which is exactly why it is used where the others would make a sauce too sweet.Sweet, fruity and faintly bitter, with only a mild warmth behind it. The bitterness is part of the character rather than a defect — it is what stops ancho reading as jam — and it is why toasting is a short operation: a scorched ancho becomes bitter in a way that cannot be balanced out.
AromaNutty and woody with a light smokiness, and noticeably less sweet than ancho or mulato. Toasting brings the nuttiness forward sharply and is the reason it is almost always toasted.Raisin and prune with a faint tobacco edge and something close to cocoa. Sweet rather than sharp, and quite unlike the aroma of any fresh chilli.

Which should you buy?

Cascabel

Buy whole, and the reason here is unusually specific: the flavour that makes cascabel worth having does not exist in the raw pod. The nuttiness develops during toasting, so a powder has already missed the step that creates it and tastes thin and vegetal by comparison. The whole pod also allows the seeds to be shaken out, which matters more for this chilli than for the thick-fleshed ones because the pod is hollow and the seeds are a large share of it — leaving them in or taking them out visibly changes the heat. And only a whole pod can be shaken, which is how anyone tells cascabel from the other round chillies sold under the same regional name.

Ancho

Buy whole pods, and buy them by feel rather than by sight: a bag that folds is worth more than one that looks handsome and snaps. They are cheap, they freeze well, and the two-minute toast-and-soak is the whole technique. Ground ancho is a reasonable thing to own for rubs and a poor substitute for a rehydrated pod, because a large part of what ancho contributes to a sauce is body rather than flavour — a mole made with powder is thin in a way no amount of extra powder fixes.

Full detail on cascabel and ancho, including evidence grades and sources for every claim above.